TLDR: always run Modify->Compute All in Fusion before exporting a STEP or STL.
From time to time I have used Modify->Compute All in Fusion to update the model if something is obviously not applied properly. But normally I don’t use Compute All because things generally stay the same.
I had a feature related to adding a tolerance to fit a 12 small identical parts into a large part. It printed fine in previous prints when I printed a small subsection of the part for testing and I didn’t change anything about that feature since, I just changed some other unrelated features. Then yesterday I printed the full part, it was a 6 hour print and when I tried to insert the small parts they were too tight and wouldn’t fit.
I measured the slot on Orca and in Fusion and it was the same, but when I measured the small part, there was no tolerance. I confirmed that the feature wasn’t suppressed and that the timeline wasn’t set in history before the feature. I then ran a Compute All and it corrected the part.
So the lesson I learned is to always run Compute All, at least for a larger time and filament wasting print.